Instructions to Authors

Cytologia, an international journal of cytogenetics and cell biology, will receive original articles in the field of cell sciences, including cytology, cytogenetics, and cell biology in a wide range of organisms using a variety of research methods. The editors will decide whether manuscripts meet the publication criteria for the journal after peer review. Cytologia is published quarterly in March, June, September, and December, unless otherwise stated.

Cytologia publishes three types of manuscript
Regular Articles describe new, original work and should be arranged in the following sequence: (1) Title page, (2) Text comprising the Introduction, Materials and methods, Results, and Discussion, (3) Acknowledgments, (4) References, (5) Tables, (6) Figure legends, and (7) Figures. If articles exceed SIX printed pages, overage fees will be charged.
Cytologia Focus papers are designed to be mini reviews that highlight recent findings, new concepts, or an overview of Cytologia articles. This article should have only one figure. Preparation of the manuscript follows the same procedure as for Regular Articles. The sections of a Cytologia Focus paper should be arranged in the following sequence: (1) Title page, (2) Text (maximum 4,000 words including spaces), (3) Acknowledgments, (4) References, and (5) Color Figure.
Technical Notes aim to introduce new techniques and findings with a cover color photo. Usually, one of the editors will invite authors to contribute to this type of manuscript; however, submission by the authors will also be considered. Authors with novel topics, including techniques, and impressive figures are encouraged to make contact with the editorial office. The sections of a Technical Note should be arranged in the following sequence: (1) Title page, (2) Text (within 1,000 words including spaces, acknowledgments, and references), and (3) a color figure as a cover photo.

1. Submission of manuscripts

Manuscripts should be submitted online using the Editorial System for Cytologia
( Before your submission, we recommend that you obtain an ORCID ID ( It is important that the submitted manuscript is written in grammatically correct English and the editors strongly recommended that articles are checked by an English editorial service or a colleague who is a native speaker of English and submitted in a complete and finished form.

  1. All manuscripts should be submitted through the Editorial System for Cytologia
  2. Upon registration, please respond to all questions.
  3. Upload files as “Submission file”
    Item “*Manuscript”:Manuscript file including all tables and figures in an MS word file (.doc or .docx). Please turn “Track Changes” off. This file will be converted by the editorial system to a PDF file for the peer reviewers. Manuscripts must contain a title page, manuscript, tables, and figures. Please check that the PDF file built by the editorial system is complete and contains all relevant information. Next, upload tables and figures as “Table,” “Figure,” or “Color figure” for printing.
    Item “Table”: Tables can be upload in MS word files (.doc or .docx) or Excel files (.xls or .xlsx).
    Item “Figure” and “Color figure”: Figure files can be uploaded in JPG, TIFF, EPS, PPT or PSD format. Upload regular figures as Item “Figure” and figures that needto be printed in color as Item “Color figure”.
    Item “Supplement”: Supplementary data can be uploaded here. Supplement files are not printed and supplied on the Cytologia website. (

Should the corresponding author not receive a confirmation by e-mail that the manuscript has been received by the Editorial Office, s/he should check the web submission and make an inquiry to the following office.

Masahiro HIZUME
The Editor-in-Chief of Cytologia
The Japan Mendel Society, 
Epoch Hongo, Hongo 2-27-2,
Bunkyo-ku, Tokyo, 113-0033 Japan 
Fax: +81-3-3814-5352

2. Preparation of manuscripts

  1. Manuscripts must be written in English and prepared using MS word. It is preferable to use a normal, plain font (e.g., 11 point Times New Roman) and 20 point line spacing or double spaced lines on A4 size pages with 3 cm margins. Page numbers and continuous line numbers should be used throughout the manuscript. Before submission, the ”Track Changes” function should be turned off.
  2. Sections should be arranged in the following sequence: (1) Title page, (2) Summary, (3) Keywords, (4) Text comprising the Introduction, Materials and methods, Results, and Discussion (for Technical Notes and Cytologia Focus papers, the text does not need sectioning), (5) Acknowledgments, (6) References, (7) Tables, (8) Figure legends, and (9) Figures and Color Figures.
  3. The title page should consist of: (a) the title of the article, (b) the full names of all authors, (c) affiliations with appropriate addresses, (d) footnotes containing the e-mail addresses of the corresponding authors (if any) and their current addresses, (e) the number of tables, figures, and color figures, and (f) a running title with less than 80 characters (including spaces). (g) Where there are multiple authors, a superscript asterisk should be added to the end of the name of the corresponding author. Authors from different institutions should be identified using a numerical superscript at the end of their name and beginning of their affiliations. (h) If your manuscript has already been opened as a preprint in the servers, including bioRxiv and ASAPbio, you should describe URL of the preprint repository.
  4. A summary should state the scope of the work and the principal findings in no more than 250 words. References should not be included.
  5. Keywords (not more than 6 words or phrases) should be provided after the summary.
  6. The text should be clear and concise under the headings of : Introduction, Materials and methods, Results, and Discussion. This should be followed by Acknowledgments, References, Tables, Figure legends, and Figures.
  7. Units of measurement should be in SI (Système International d'Unités).
  8. In-text citations should be made by the authors’ names (for three or more authors, use et al., e.g., Kuroiwa et al.) followed by the year of publication.
    At the head of a sentence: Darlington and Wylie (1955) reported.
    In the case of more than three authors: Kuroiwa et al. (1981a) indicated.
    At the end of a sentence: (Darlington and Wylie 1955, Kuroiwa et al. 1981a, Kuroiwa 1989).
  9. References should be arranged alphabetically according to the names of author(s). Each reference should be given in the following form: A reference written by more than 20 authors should be written as the first author name followed by et al.
    • Book:
      Darlington, C. D. and Wylie, A. P. 1955. Chromosome Atlas of Flowering Plants. Allen and Unwin Ltd., London.
    • Book chapter:
      Bennett, M. D. 1983. The Spatial Distribution of Chromosomes. In: Brandham, P. E. and Bennett, M. D. (eds.). Kew Chromosome Conference II. Allen and Urwin Ltd., London. pp. 71-90.
    • Journal article: The abbreviation of journal name should follow the style used in Journals in the NCBI Databases (
      Kuroiwa, T., Nishibayashi, S., Kawano, S. and Suzuki, S. 1981a. Visualization of DNA in various phages (T4, X, T7, ø29) by ethidium bromide epi-fluorescent microscopy. Experientia 37: 969–971.
      Kuroiwa, T., Suzuki, S., Ogawa, K. and Kawano, S. 1981b. The chloroplast nucleus: distribution, number, size and shape, and a model for the multiplication of the chloroplast genome during chloroplast development. Plant Cell Physiol. 22: 381–396.
    • Conference proceedings: 
      Zachleder, V. and Kawano, S. 2012. Comparison of starch and oil production by Chlamydomonas reinhardtii and other algae. In: The 15th International Conference on the Cell and Molecular Biology of Chlamydomonas, Potsdam. p.184.
    • Non-English references: References in languages other than English, German, Spanish and French must be translated into English followed by the original language in parenthesis :
      Ota, S. and Kawano, S. 2017. Chlorella biomaterials: phosphate, starch, oil and possible involvement of autophagy. Plant Morphol. 29: 57-61. (in Japanese)
    • For Web sites: Direct quotation of websites is not accepted.
  10. Each table should be provided on a separate sheet and presentedd after the references. Tables should be provided with headings on the same page. Vertical lines should not be used in tables.
  11. Illustration and images should be prepared high resolution using at least 300 dpi in print (Full width is 2008 pixels and half is 945 pixels) in JPG, TIF, EPS, PSD or PPT format. Never use MS word file (.doc or .docx). The magnifications of figures must be indicated with a scale bars within the images. Letterings should be large enough to permit reduction of printing.
  12. Color figures can be printed provided that the author pays the additional cost (see section 5). Color figure files should be uploaded as Item “Color figure”.

3. Proofreading

A proof is sent only once to the corresponding author. Proofreading should be limited to the correction of typographical errors. The setting costs incurred by excess changes will be charged to the author.

4. Author's charges

The author is requested to pay extra costs in the following cases:

  1. The author is supposed to pay a publication fee of US $40 (4,000 yen) for a publishing manuscript. (See section 7 for methods of payment.)
  2. For the additional pages, i.e., exceeding SIX printed pages, the author is required to pay US $100 (10,000 yen) per each additional page. Changes made at the proofreading stage, except for the correction of typographical errors, are charged to the author.
  3. If the first author or the corresponding author is a member of The Japan Mendel Society, the publication fee won’t be charged up to four manuscripts per year per person.

5. Color Charge

If the manuscript contains one or more color illustrations and/or figures, a color fee of US $200 (20,000 yen) for each color page will be charged to the author.

6. Reprints

Reprints provide authors with a means of obtaining wider distribution for their work. A reprint order request must be sent together with the proof to the editorial office. It must be received before the Journal goes to press. Do not delay ordering your reprints.

Delivery and invoices: Reprints are normally shipped 3 weeks after publication of the Journal. Invoices are emailed to the author before the time of shipment. In the case of orders chargeable to institutions, an official purchase order must be raised before the reprint shipment can be released.

The prices in US dollars quoted here are for reprints ordered by and paid for by the author, or by a sponsoring non-profit organization, and only apply if orders are in our possession before the journal goes to press. There is a minimum order 20 copies; quantities above 50 copies will be charged for in multiples of 50 copies.

Pages Per 20 copies Per 50 copies Per 100 copies
1-4 US $50 (5,000 yen) US $100 (10,000 yen) US $200 (20,000 yen)
5-6 US $60 (6,000 yen) US $120 (12,000 yen) US $240 (24,000 yen)
7-8 US $70 (7,000 yen) US $140 (14,000 yen) US $280 (28,000 yen)
9-10 US $80 (8,000 yen) US $160 (16,000 yen) US $320 (32,000 yen)
11-12 US $90 (9,000 yen) US $180 (18,000 yen) US $360 (36,000 yen)

7. Methods of Payment

Authors should pay the “publication fee,” “page charge,” “color charge,” and “reprints charge” in advance. Payment may be made using the following means. Accepted credit cards (VISA and MASTERCARD) can be used for paying annual JMS membership fees and publication fees (for non-members of the society), as well as page, color, and reprint charges

Please contact “” for information regarding your payment by credit card. We will send you the web address (URL) where you can complete your credit card payment.

(June 21, 2021, renewed)