Instructions to Authors

CYTOLOGIA, International Journal of Cytogenetics and Cell Biology, will receive original articles in the field of Cell Sciences including cytology, cytogenetics, and cell biology in a wide range of organisms using a variety of research methods. Editors will decide whether manuscripts meet publication criteria in this journal after peer review. CYTOLOGIA is published quarterly in March, June, September and December except otherwise stated.

Cytologia has three types of manuscripts,
Regular Article describes new, original work and should be arranged in the following sequence: (1) Title page, (2) Text composing of Introduction, Materials and methods, Results, and Discussion, (3) Acknowledgments, (4) References, (5) Tables, (6) Figure legends, and (7) Figures. Articles should be no more than SIX printed pages.
Cytologia Focus is designed to be a mini review that high-lights of recent findings, new concepts or an overview of CYTOLOGIA articles. This article should have with only one figure. Preparation of the manuscript is the same as Regular Articles. Sections of the Cytologia Focus should be arranged in the following sequence: (1) Title page, (2) Text (within 4,000 words including spaces), (3) Acknowledgments, (4) References, and (5) Color Figure. The articles should comprise no more FIVE printed pages.
Technical Note aims to introduce new techniques and findings with a cover color photo. Usually, either of editors will invite authors to contribute this type of manuscripts; however, submission by the authors will also be considered. Authors with novel topics including techniques and impressive figures are encouraged to have contact with the editorial office. Sections of the Technical Note should be arranged in following sequence: (1) Title page, (2) Text (within 1,000 words including spaces, acknowledgments, and references), and (3) Color Figure as a cover photo.

1. Submission of manuscripts

Manuscripts should be submitted on line using the Editorial System for CYTOLOGIA
. Before your submission, we recommend that you get ORCID ID ( It is important that the submitted manuscript is written in grammatically correct English and the editors recommended strongly that articles are checked by an English editorial service or a colleague who is a native speaker of English and submitted in a complete and finished form.

  1. All manuscripts should be submitted through the Editorial System for CYTOLOGIA
  2. Upon registration, please respond to initial all questions.
  3. Upload files as “Submission file”
    Item “*Manuscript”: Manuscript file including all tables and figures in a MS word file (doc or docx). Please turn “Track Changes” off. This file will be converted by the editorial system to PDF file for peer reviewers. Manuscript must contain Title page, Manuscript, Tables and Figures. Please check PDF file build by the editorial system to be complete and contain all information.
    Next, upload also tables and figures as “Table” “Figure” “Color figure” for printing.
    Item “Table”: Tables can be upload in MS word files (doc or docx) or Excel files (xls or xlsx).
    Item “Figure” and “Color figure”: Figure files can be uploaded in JPG, TIFF, EPS, PPT or PSD format. Upload regular figures as Item “Figure”and figures needing print in color as Item “Color figure”.
    Item “Supplement”: Supplement data can be upload here. Supplement files are not printed and supplied on the web site of Cytologia (

Should the corresponding author not receive a confirmation by e-mail that the manuscript has been received by the Editorial Office, they should check their web submission and inquire to the following editorial office.

Masahiro HIZUME
The Editor-in-Chief of CYTOLOGIA
The Japan Mendel Society, 
Esperanza V Bldg., Hongo 2-27-2,
Bunkyo-ku, Tokyo, 113-0033 Japan 
Fax: +81-3-3814-5352

2. Preparation of manuscripts

  1. Manuscripts must be written in English prepared using the MS word. It is preferable to use a normal, plain font (e.g. 11 point Times New Roman) and 20 lines or double space lines in A4 size paper with 3-cm margins. The page numbers and continues line numbers should be used through the manuscript. Before submission “Track Changes” function should be turned off.
  2. Sections should be arranged in the following sequence: (1) Title page, (2) Summary, (3) Keywords, (4)Text composing introduction, Materials and methods, Results, and Discussion, (In the case of Technical Note and Cytologia Focus, Text does not need sectioning.), (5) Acknowledgements, (6) References, (7) Tables, (8) Figure legends,(9) Figures and Color Figures.
  3. Title page should comprise of (a) title of the article, (b) full names of all authors, (c) affiliations with appropriate addresses, (d) footnotes of e-mail address of the corresponding author, if any, present address, etc., (e) numbers of Tables, Figures and Color Figures, (f) running title with less than 80 characters that include spaces. (g) Where there are multiple authors, a superscript asterisk should be added to the end of the name of the corresponding author. Authors with different institutions should be identified using numerical superscript at the end of their name and beginning of their affiliations.
  4. Summary should state the scope of the work and the principal findings in no more than 250 words. References should not be included.
  5. Keywords (not more than 6 words or phrases) should be written after the summary.
  6. Text should be clear and concise under the headings: Introduction, Materials and methods, Results, and Discussion followed by Acknowledgments, References, Tables, Figure legends, and Figures.
  7. Units of measurement should be in SI (Système International d'Unités) as a rule.
  8. Text references should be made by the authors' names (for three or more authors, use et al., e.g. Kuroiwa et al.) followed by the year of publication.
    At the head of sentence: Darlington and Wylie (1955) reported.
    In the case of more than three authors: Kuroiwa et al. (1981a) indicated.
    At the end: (Darlington and Wylie 1955, Kuroiwa et al. 1981a, Kuroiwa 1989).
  9. References should be arranged alphabetically according to the names of author(s). Each reference should be given in the following form: The reference written by more than 20 authors should be written as the first author name et al.
    • Book:
      Darlington, C. D. and Wylie, A. P. 1955. Chromosome Atlas of Flowering Plants. Allen and Unwin Ltd., London.
    • Book chapter:
      Bennett, M. D. 1983. The Spatial Distribution of Chromosomes. In: Brandham, P. E. and Bennett, M. D. (eds.). Kew Chromosome Conference II. Allen and Urwin Ltd., London. pp. 71-90.
    • Journal article: Abbreviation of journal name should be followed in Journals in NCBI Databases ( for example:
      Kuroiwa, T., Nishibayashi, S., Kawano, S. and Suzuki, S. 1981a. Visualization of DNA in various phages (T4, X, T7, ø29) by ethidium bromide epi-fluorescent microscopy. Experientia 37: 969–971.
      Kuroiwa, T., Suzuki, S., Ogawa, K. and Kawano, S. 1981b. The chloroplast nucleus: distribution, number, size and shape, and a model for the multiplication of the chloroplast genome during chloroplast development. Plant Cell Physiol. 22: 381–396.
    • Conference proceedings: 
      Zachleder, V. and Kawano, S. 2012. Comparison of starch and oil production by Chlamydomonas reinhardtii and other algae. In: The 15th International Conference on the Cell and Molecular Biology of Chlamydomonas, Potsdam. p.184.
    • For non-English references: References in languages other than English, German, Spanish and French must be translated into English followed by the original language in parenthesis for example:
      Ota, S. and Kawano, S. 2017. Chlorella biomaterials: phosphate, starch, oil and possible involvement of autophagy. Plant Morphol. 29: 57-61. (in Japanese)
    • For Web sites: Quotation of websites is not admitted.
  10. Each table should be provided on a separate sheet and collected after the references. Tables should be provided with headings on the same sheet. Vertical lines should not be used in tables.
  11. Figures of illustrations and images should be prepared high resolution using at least 300 dpi in print (Full width is 2008 pixels and half is 945 pixels) in JPG, TIF, EPS, PSD or PPT format. Never use MS word file (doc or docx). Magnifications of figures must be indicated with scale bars in the images. Letterings should be large enough to permit reduction of printing.
  12. Color figures can be printed provided that the author pays the additional cost (see section 5). Color figure files should be uploaded as Item “Color figure”.

3. Proofreading

A proof is sent just one time to the author. Proofreading should be limited to the correction of typographical errors. The setting costs incurred by excess alterations will be charged to the author.

4. Author's charges

The author is requested to pay extra costs in the following cases:

  1. The author is supposed to pay a publication fee of US $40 (4,000 yen) for a publishing manuscript. (See section 7 for methods of payment.)
  2. For the additional pages exceeding SIX printed pages, the author is required to pay US $100 (10,000 yen) per each additional page. The changes made at the proofreading stage except for the correction of typographical errors are charged to the author.
  3. If the first author or the corresponding author is a member of The Japan Mendel Society, the publication fee won’t be charged up to four manuscripts for one year per person.

5. Color Charge

If the manuscript contains one or more color illustrations and/or figures, the color fee of US $200 (20,000 yen) for each color page will be charged to the author.

6. Reprints

Reprints provide authors with a means of obtaining wider distribution for their work. A reprint order request must be sent together with the proof to the editorial office. It must be received before the Journal goes to press. Do not delay ordering your reprints.

Delivery and invoices: Reprints are normally shipped 3 weeks after publication of the Journal. Invoices are emailed the author before the time of shipment. In the case of orders chargeable to Institutions, an official Purchase Order must be in hand before the reprint shipment can be released.

The prices in US dollars quoted herein are for reprints ordered by and paid for by the author only, or by sponsoring non-profit organization, and apply only if orders are in our possession before the Journal goes to press. Minimum order 20 copies; quantities above 50 copies will be charged for in multiples of 50 copies.

Pages Per 20 copies Per 50 copies Per 100 copies
1-4 US $50 (5,000 yen) US $100 (10,000 yen) US $200 (20,000 yen)
5-6 US $60 (6,000 yen) US $120 (12,000 yen) US $240 (24,000 yen)
7-8 US $70 (7,000 yen) US $140 (14,000 yen) US $280 (28,000 yen)
9-10 US $80 (8,000 yen) US $160 (16,000 yen) US $320 (32,000 yen)
11-12 US $90 (9,000 yen) US $180 (18,000 yen) US $360 (36,000 yen)

7. Methods of Payment

Authors should pay in advance the dues of “Publication fee” “Over page charge” “Color charge” and “Reprints”. Author’s dues may be paid using the following means. Accepted credit cards (VISA and MASTER) are used for paying annual fees of the Japan Mendel Society, and publication fees (for non-members of the society), page charges, reprints etc. of CYTOLOGIA, as extra charges are usually not requested.

Please contact “” for information of your payment by accepted credit cards. We will send you the Web address (URL) where you can complete your credit card payment.

(February 29, 2020, renewed)